query	query_gene_id	query_protein_id	source_reference	hit_rank	swissprot_acc	swissprot_entry	database_section	target_gene_id	description	organism	homolog_gene_name	percent_identity	alignment_length	query_coverage	evalue	bitscore	confidence	evidence_type	source_db	reference_gene_id	reference_function_zh	reference_function_en	reference_papers_json
LXL04_022485	gene-LXL04_022485	KAL4575634.1	OH-1004	2	Q9C5U0	AHK4_ARATH	Swiss-Prot		Histidine kinase 4 OS=Arabidopsis thaliana OX=3702 GN=AHK4 PE=1 SV=1	Arabidopsis thaliana	AHK4	68.937	969	97.065	0.0	1334.0	strong_database_homology	BLASTP to reviewed UniProt Swiss-Prot	UniProt Swiss-Prot current_release 2026-06-10	At2g01830	与细菌双组分调节剂相关的细胞分裂素 受体。	Cytokinins (CK) receptor related to bacterial two-component regulators.	"[{""pmid"":""11230563"",""title"":""The Arabidopsis sensor His-kinase, AHk4, can respond to cytokinins."",""doi"":""10.1093/pcp/pce037"",""url"":""https://pubmed.ncbi.nlm.nih.gov/11230563/"",""year"":""2001"",""journal"":""Plant & cell physiology"",""authors"":""Suzuki T, Miwa K, Ishikawa K, Yamada H, Aiba H, Mizuno T."",""matched_aliases"":[""AHK4""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""11234017"",""title"":""Identification of CRE1 as a cytokinin receptor from Arabidopsis."",""doi"":""10.1038/35059117"",""url"":""https://pubmed.ncbi.nlm.nih.gov/11234017/"",""year"":""2001"",""journal"":""Nature"",""authors"":""Inoue T, Higuchi M, Hashimoto Y, Seki M, Kobayashi M, Kato T, Tabata S, Shinozaki K, Kakimoto T."",""matched_aliases"":[""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""11479382"",""title"":""The AHK4 gene involved in the cytokinin-signaling pathway as a direct receptor molecule in Arabidopsis thaliana."",""doi"":""10.1093/pcp/pce094"",""url"":""https://pubmed.ncbi.nlm.nih.gov/11479382/"",""year"":""2001"",""journal"":""Plant & cell physiology"",""authors"":""Ueguchi C, Sato S, Kato T, Tabata S."",""matched_aliases"":[""AHK4""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""11577198"",""title"":""The Arabidopsis AHK4 histidine kinase is a cytokinin-binding receptor that transduces cytokinin signals across the membrane."",""doi"":""10.1093/pcp/pce127"",""url"":""https://pubmed.ncbi.nlm.nih.gov/11577198/"",""year"":""2001"",""journal"":""Plant & cell physiology"",""authors"":""Yamada H, Suzuki T, Terada K, Takei K, Ishikawa K, Miwa K, Yamashino T, Mizuno T."",""matched_aliases"":[""AHK4"",""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""12354925"",""title"":""Characterization of the ARR15 and ARR16 response regulators with special reference to the cytokinin signaling pathway mediated by the AHK4 histidine kinase in roots of Arabidopsis thaliana."",""doi"":""10.1093/pcp/pcf121"",""url"":""https://pubmed.ncbi.nlm.nih.gov/12354925/"",""year"":""2002"",""journal"":""Plant & cell physiology"",""authors"":""Kiba T, Yamada H, Mizuno T."",""matched_aliases"":[""AHK4"",""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""12410813"",""title"":""Mutations at CRE1 impair cytokinin-induced repression of phosphate starvation responses in Arabidopsis."",""doi"":""10.1046/j.1365-313x.2002.01431.x"",""url"":""https://pubmed.ncbi.nlm.nih.gov/12410813/"",""year"":""2002"",""journal"":""The Plant journal : for cell and molecular biology"",""authors"":""Franco-Zorrilla JM, Martin AC, Solano R, Rubio V, Leyva A, Paz-Ares J."",""matched_aliases"":[""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""15053761"",""title"":""Interallelic complementation at the Arabidopsis CRE1 locus uncovers independent pathways for the proliferation of vascular initials and canonical cytokinin signalling."",""doi"":""10.1111/j.1365-313x.2004.02023.x"",""url"":""https://pubmed.ncbi.nlm.nih.gov/15053761/"",""year"":""2004"",""journal"":""The Plant journal : for cell and molecular biology"",""authors"":""de León BG, Zorrilla JM, Rubio V, Dahiya P, Paz-Ares J, Leyva A."",""matched_aliases"":[""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""15144379"",""title"":""A novel regulatory pathway of sulfate uptake in Arabidopsis roots: implication of CRE1/WOL/AHK4-mediated cytokinin-dependent regulation."",""doi"":""10.1111/j.1365-313x.2004.02079.x"",""url"":""https://pubmed.ncbi.nlm.nih.gov/15144379/"",""year"":""2004"",""journal"":""The Plant journal : for cell and molecular biology"",""authors"":""Maruyama-Nakashita A, Nakamura Y, Yamaya T, Takahashi H."",""matched_aliases"":[""AHK4"",""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""15155880"",""title"":""Histidine kinase homologs that act as cytokinin receptors possess overlapping functions in the regulation of shoot and root growth in Arabidopsis."",""doi"":""10.1105/tpc.021477"",""url"":""https://pubmed.ncbi.nlm.nih.gov/15155880/"",""year"":""2004"",""journal"":""The Plant cell"",""authors"":""Nishimura C, Ohashi Y, Sato S, Kato T, Tabata S, Ueguchi C."",""matched_aliases"":[""AHK4"",""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""15166290"",""title"":""In planta functions of the Arabidopsis cytokinin receptor family."",""doi"":""10.1073/pnas.0402887101"",""url"":""https://pubmed.ncbi.nlm.nih.gov/15166290/"",""year"":""2004"",""journal"":""Proceedings of the National Academy of Sciences of the United States of America"",""authors"":""Higuchi M, Pischke MS, Mähönen AP, Miyawaki K, Hashimoto Y, Seki M, Kobayashi M, Shinozaki K, Kato T, Tabata S, Helariutta Y, Sussman MR, Kakimoto T."",""matched_aliases"":[""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""15509853"",""title"":""Two cytokinin receptors of Arabidopsis thaliana, CRE1/AHK4 and AHK3, differ in their ligand specificity in a bacterial assay."",""doi"":""10.1093/pcp/pch132"",""url"":""https://pubmed.ncbi.nlm.nih.gov/15509853/"",""year"":""2004"",""journal"":""Plant & cell physiology"",""authors"":""Spíchal L, Rakova NY, Riefler M, Mizuno T, Romanov GA, Strnad M, Schmülling T."",""matched_aliases"":[""AHK4"",""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""15728338"",""title"":""Topolins and hydroxylated thidiazuron derivatives are substrates of cytokinin O-glucosyltransferase with position specificity related to receptor recognition."",""doi"":""10.1104/pp.104.057174"",""url"":""https://pubmed.ncbi.nlm.nih.gov/15728338/"",""year"":""2005"",""journal"":""Plant physiology"",""authors"":""Mok MC, Martin RC, Dobrev PI, Vanková R, Ho PS, Yonekura-Sakakibara K, Sakakibara H, Mok DW."",""matched_aliases"":[""AHK4"",""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""15923327"",""title"":""Interaction between phosphate-starvation, sugar, and cytokinin signaling in Arabidopsis and the roles of cytokinin receptors CRE1/AHK4 and AHK3."",""doi"":""10.1104/pp.105.060517"",""url"":""https://pubmed.ncbi.nlm.nih.gov/15923327/"",""year"":""2005"",""journal"":""Plant physiology"",""authors"":""Franco-Zorrilla JM, Martín AC, Leyva A, Paz-Ares J."",""matched_aliases"":[""AHK4"",""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""16246292"",""title"":""A live cell hormone-binding assay on transgenic bacteria expressing a eukaryotic receptor protein."",""doi"":""10.1016/j.ab.2005.09.012"",""url"":""https://pubmed.ncbi.nlm.nih.gov/16246292/"",""year"":""2005"",""journal"":""Analytical biochemistry"",""authors"":""Romanov GA, Spíchal L, Lomin SN, Strnad M, Schmülling T."",""matched_aliases"":[""AHK4"",""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""16357038"",""title"":""Cytokinin receptors are required for normal development of auxin-transporting vascular tissues in the hypocotyl but not in adventitious roots."",""doi"":""10.1093/pcp/pci240"",""url"":""https://pubmed.ncbi.nlm.nih.gov/16357038/"",""year"":""2006"",""journal"":""Plant & cell physiology"",""authors"":""Kuroha T, Ueguchi C, Sakakibara H, Satoh S."",""matched_aliases"":[""AHK4""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""16361392"",""title"":""Arabidopsis cytokinin receptor mutants reveal functions in shoot growth, leaf senescence, seed size, germination, root development, and cytokinin metabolism."",""doi"":""10.1105/tpc.105.037796"",""url"":""https://pubmed.ncbi.nlm.nih.gov/16361392/"",""year"":""2006"",""journal"":""The Plant cell"",""authors"":""Riefler M, Novak O, Strnad M, Schmülling T."",""matched_aliases"":[""AHK4"",""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""16753566"",""title"":""Cytokinins regulate a bidirectional phosphorelay network in Arabidopsis."",""doi"":""10.1016/j.cub.2006.04.030"",""url"":""https://pubmed.ncbi.nlm.nih.gov/16753566/"",""year"":""2006"",""journal"":""Current biology : CB"",""authors"":""Mähönen AP, Higuchi M, Törmäkangas K, Miyawaki K, Pischke MS, Sussman MR, Helariutta Y, Kakimoto T."",""matched_aliases"":[""AHK4"",""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""17075078"",""title"":""Biochemical characteristics and ligand-binding properties of Arabidopsis cytokinin receptor AHK3 compared to CRE1/AHK4 as revealed by a direct binding assay."",""doi"":""10.1093/jxb/erl179"",""url"":""https://pubmed.ncbi.nlm.nih.gov/17075078/"",""year"":""2006"",""journal"":""Journal of experimental botany"",""authors"":""Romanov GA, Lomin SN, Schmülling T."",""matched_aliases"":[""AHK4"",""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""17216481"",""title"":""The floral volatile, methyl benzoate, from snapdragon (Antirrhinum majus) triggers phytotoxic effects in Arabidopsis thaliana."",""doi"":""10.1007/s00425-006-0464-0"",""url"":""https://pubmed.ncbi.nlm.nih.gov/17216481/"",""year"":""2007"",""journal"":""Planta"",""authors"":""Horiuchi J, Badri DV, Kimball BA, Negre F, Dudareva N, Paschke MW, Vivanco JM."",""matched_aliases"":[""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""17956858"",""title"":""Identification of amino acid substitutions that render the Arabidopsis cytokinin receptor histidine kinase AHK4 constitutively active."",""doi"":""10.1093/pcp/pcm145"",""url"":""https://pubmed.ncbi.nlm.nih.gov/17956858/"",""year"":""2007"",""journal"":""Plant & cell physiology"",""authors"":""Miwa K, Ishikawa K, Terada K, Yamada H, Suzuki T, Yamashino T, Mizuno T."",""matched_aliases"":[""AHK4"",""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""17965178"",""title"":""Cytokinin receptors are involved in alkamide regulation of root and shoot development in Arabidopsis."",""doi"":""10.1104/pp.107.107953"",""url"":""https://pubmed.ncbi.nlm.nih.gov/17965178/"",""year"":""2007"",""journal"":""Plant physiology"",""authors"":""López-Bucio J, Millán-Godínez M, Méndez-Bravo A, Morquecho-Contreras A, Ramírez-Chávez E, Molina-Torres J, Pérez-Torres A, Higuchi M, Kakimoto T, Herrera-Estrella L."",""matched_aliases"":[""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""18077346"",""title"":""Functional analysis of AHK1/ATHK1 and cytokinin receptor histidine kinases in response to abscisic acid, drought, and salt stress in Arabidopsis."",""doi"":""10.1073/pnas.0706547105"",""url"":""https://pubmed.ncbi.nlm.nih.gov/18077346/"",""year"":""2007"",""journal"":""Proceedings of the National Academy of Sciences of the United States of America"",""authors"":""Tran LS, Urao T, Qin F, Maruyama K, Kakimoto T, Shinozaki K, Yamaguchi-Shinozaki K."",""matched_aliases"":[""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""18397377"",""title"":""Cytokinins negatively regulate the root iron uptake machinery in Arabidopsis through a growth-dependent pathway."",""doi"":""10.1111/j.1365-313x.2008.03502.x"",""url"":""https://pubmed.ncbi.nlm.nih.gov/18397377/"",""year"":""2008"",""journal"":""The Plant journal : for cell and molecular biology"",""authors"":""Séguéla M, Briat JF, Vert G, Curie C."",""matched_aliases"":[""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""18571199"",""title"":""Mechanism-based inhibitors of cytokinin oxidase/dehydrogenase attack FAD cofactor."",""doi"":""10.1016/j.jmb.2008.05.044"",""url"":""https://pubmed.ncbi.nlm.nih.gov/18571199/"",""year"":""2008"",""journal"":""Journal of molecular biology"",""authors"":""Kopecný D, Sebela M, Briozzo P, Spíchal L, Houba-Hérin N, Masek V, Joly N, Madzak C, Anzenbacher P, Laloue M."",""matched_aliases"":[""AHK4""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""18785832"",""title"":""The role of auxins and cytokinins in the mutualistic interaction between Arabidopsis and Piriformospora indica."",""doi"":""10.1094/mpmi-21-10-1371"",""url"":""https://pubmed.ncbi.nlm.nih.gov/18785832/"",""year"":""2008"",""journal"":""Molecular plant-microbe interactions : MPMI"",""authors"":""Vadassery J, Ritter C, Venus Y, Camehl I, Varma A, Shahollari B, Novák O, Strnad M, Ludwig-Müller J, Oelmüller R."",""matched_aliases"":[""CRE1""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""19032596"",""title"":""The purine derivative PI-55 blocks cytokinin action via receptor inhibition."",""doi"":""10.1111/j.1742-4658.2008.06777.x"",""url"":""https://pubmed.ncbi.nlm.nih.gov/19032596/"",""year"":""2009"",""journal"":""The FEBS journal"",""authors"":""Spíchal L, Werner T, Popa I, Riefler M, Schmülling T, Strnad M."",""matched_aliases"":[""AHK4"",""CRE1""],""evidence"":""UniProt FUNCTION; 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LXL04_022485	gene-LXL04_022485	KAL4575634.1	OH-1004	5	Q9C5U2	AHK2_ARATH	Swiss-Prot		Histidine kinase 2 OS=Arabidopsis thaliana OX=3702 GN=AHK2 PE=1 SV=1	Arabidopsis thaliana	AHK2	54.425	983	97.484	0.0	994.0	strong_database_homology	BLASTP to reviewed UniProt Swiss-Prot	UniProt Swiss-Prot current_release 2026-06-10	At5g35750	与细菌双组分调节剂相关的细胞分裂素 受体。	Cytokinins (CK) receptor related to bacterial two-component regulators.	"[{""pmid"":""15155880"",""title"":""Histidine kinase homologs that act as cytokinin receptors possess overlapping functions in the regulation of shoot and root growth in Arabidopsis."",""doi"":""10.1105/tpc.021477"",""url"":""https://pubmed.ncbi.nlm.nih.gov/15155880/"",""year"":""2004"",""journal"":""The Plant cell"",""authors"":""Nishimura C, Ohashi Y, Sato S, Kato T, Tabata S, Ueguchi C."",""matched_aliases"":[""AHK2""],""evidence"":""UniProt FUNCTION; UniProt/PubMed metadata and gene/species abstract checks; EuropePMC cross-check where available"",""metadata_sources"":[""UniProt"",""PubMed"",""EuropePMC""],""fulltext_reviewed"":false},{""pmid"":""15166290"",""title"":""In planta functions of the Arabidopsis cytokinin receptor family."",""doi"":""10.1073/pnas.0402887101"",""url"":""https://pubmed.ncbi.nlm.nih.gov/15166290/"",""year"":""2004"",""journal"":""Proceedings of the National Academy of Sciences of the United States of America"",""authors"":""Higuchi M, Pischke MS, Mähönen AP, Miyawaki K, Hashimoto Y, Seki M, Kobayashi M, Shinozaki K, Kato T, Tabata S, Helariutta Y, Sussman MR, Kakimoto T."",""matched_aliases"":[""AHK2""],""evidence"":""UniProt FUNCTION; 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LXL04_022485	gene-LXL04_022485	KAL4575634.1	OH-1004	3	A1A698	OHK4_ORYSJ	Swiss-Prot		Probable histidine kinase 4 OS=Oryza sativa subsp. japonica OX=39947 GN=HK4 PE=2 SV=1	Oryza sativa subsp. japonica	HK4	60.774	956	94.34	0.0	1133.0	strong_database_homology	BLASTP to reviewed UniProt Swiss-Prot	UniProt Swiss-Prot current_release 2026-06-10	Os03g0717700	与细菌双组分调控器相关的细胞分裂素受体。	Cytokinin receptor related to bacterial two-component regulators.	[]
LXL04_022485	gene-LXL04_022485	KAL4575634.1	OH-1004	4	A1A699	OHK6_ORYSJ	Swiss-Prot		Probable histidine kinase 6 OS=Oryza sativa subsp. japonica OX=39947 GN=HK6 PE=1 SV=1	Oryza sativa subsp. japonica	HK6	60.645	930	92.034	0.0	1113.0	strong_database_homology	BLASTP to reviewed UniProt Swiss-Prot	UniProt Swiss-Prot current_release 2026-06-10	Os02g0738400	与细菌双组分调控器相关的细胞分裂素受体。	Cytokinin receptor related to bacterial two-component regulators.	[]