Species	Reference gene ID	Possible function	Papers / DOI
Arabidopsis	At5g48390	Required for meiotic chromosome segregation.	Zip4/Spo22 is required for class I CO formation but not for synapsis completion in Arabidopsis thaliana.; 2007; DOI: 10.1371/journal.pgen.0030083; PMID: 17530928; https://pubmed.ncbi.nlm.nih.gov/17530928/ | Homologous chromosome pairing is completed in crossover defective atzip4 mutant.; 2008; DOI: 10.1016/j.bbrc.2008.03.036; PMID: 18348867; https://pubmed.ncbi.nlm.nih.gov/18348867/
Rice	Os01g0890900	Required for crossover formation, complete synapsis of homologous chromosomes and bivalent formation during meiosis.	ZIP4 in homologous chromosome synapsis and crossover formation in rice meiosis.; 2012; DOI: 10.1242/jcs.090993; PMID: 22393242; https://pubmed.ncbi.nlm.nih.gov/22393242/
Rice	UniProt: A2WXU2	Required for crossover formation, complete synapsis of homologous chromosomes and bivalent formation during meiosis.	No verified direct reference in this release