Species	Reference gene ID	Possible function	Papers / DOI
Arabidopsis	At2g41740	May act as a Ca²⁺-regulated actin-binding protein.	Arabidopsis VILLIN2 and VILLIN3 are required for the generation of thick actin filament bundles and for directional organ growth.; 2012; DOI: 10.1104/pp.111.192385; PMID: 22209875; https://pubmed.ncbi.nlm.nih.gov/22209875/ | Arabidopsis VILLIN2 and VILLIN3 act redundantly in sclerenchyma development via bundling of actin filaments.; 2012; DOI: 10.1111/j.1365-313x.2012.05044.x; PMID: 22563899; https://pubmed.ncbi.nlm.nih.gov/22563899/ | Arabidopsis villins promote actin turnover at pollen tube tips and facilitate the construction of actin collars.; 2013; DOI: 10.1105/tpc.113.110940; PMID: 23715472; https://pubmed.ncbi.nlm.nih.gov/23715472/
Arabidopsis	At3g57410	Binds actin and actin bundles independently of Ca²⁺, but severs actin filaments in a calcium-dependent manner.	Arabidopsis VILLIN1 and VILLIN3 have overlapping and distinct activities in actin bundle formation and turnover.; 2010; DOI: 10.1105/tpc.110.076240; PMID: 20807878; https://pubmed.ncbi.nlm.nih.gov/20807878/ | Arabidopsis VILLIN2 and VILLIN3 are required for the generation of thick actin filament bundles and for directional organ growth.; 2012; DOI: 10.1104/pp.111.192385; PMID: 22209875; https://pubmed.ncbi.nlm.nih.gov/22209875/ | Arabidopsis VILLIN2 and VILLIN3 act redundantly in sclerenchyma development via bundling of actin filaments.; 2012; DOI: 10.1111/j.1365-313x.2012.05044.x; PMID: 22563899; https://pubmed.ncbi.nlm.nih.gov/22563899/
Arabidopsis	At4g30160	Binds actin and actin bundles independently of Ca²⁺, but caps filament barbed ends and severs filaments in a calcium-dependent manner.	Arabidopsis VILLIN4 is involved in root hair growth through regulating actin organization in a Ca2+-dependent manner.; 2011; DOI: 10.1111/j.1469-8137.2010.03632.x; PMID: 21275995; https://pubmed.ncbi.nlm.nih.gov/21275995/ | The universal bundling activity of AtVLN4 in diffusely growing cells.; 2011; DOI: 10.4161/psb.6.9.16501; PMID: 22019634; https://pubmed.ncbi.nlm.nih.gov/22019634/
Arabidopsis	At5g57320	Major actin filament stabilizing factor and regulator of actin dynamics.	Arabidopsis VILLIN5, an actin filament bundling and severing protein, is necessary for normal pollen tube growth.; 2010; DOI: 10.1105/tpc.110.076257; PMID: 20807879; https://pubmed.ncbi.nlm.nih.gov/20807879/ | Arabidopsis villins promote actin turnover at pollen tube tips and facilitate the construction of actin collars.; 2013; DOI: 10.1105/tpc.113.110940; PMID: 23715472; https://pubmed.ncbi.nlm.nih.gov/23715472/
Rice	Os03g0356700	Ca(2+)-regulated actin-binding protein (By similarity).	VLN2 Regulates Plant Architecture by Affecting Microfilament Dynamics and Polar Auxin Transport in Rice.; 2015; DOI: 10.1105/tpc.15.00581; PMID: 26486445; https://pubmed.ncbi.nlm.nih.gov/26486445/